These include things such as environmental conditions in which the animals live, sex-specific growth, survival and reproduction through the history of the species.
The myriad factors affecting innovativeness make the historian's task paradoxically easier, by converting societal variation in innovativeness into essentially a random variable.
One possibility is that the genes and health of Eastern European Jews were affected not simply by geography but also by historical and cultural factors.
There's history here, too, as Finland was invaded by the Soviet Union in 1939 in what was named The Winter War, which happened at the beginning of World War II.
We British are as much heirs to the legacy of European culture as any other nation. Our links to the rest of Europe, the continent of Europe, have been the dominant factor in our history.