One inclusion formed somewhat later exterior to the exosporium. Another inclusion was formed within the exosporium and remained with the spore after mother cell lysis.

当芽胞形细胞裂解，晶体被释放出来，形自由的伴胞晶体，另芽胞外壁内侧形，熟被包裹芽胞外壁以内不脱落，始终和芽胞粘连起。