Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不老虎“”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员照片来测量老虎脚印长度和宽度以及脚趾角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印足迹同样是常用计算数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量印长以及趾之间角，以及10种其他变量来确定不同“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测老虎脚印长度和宽度以及脚趾之间角度，以及10种来确定不同老虎“身份”。